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Added 4 mins ago
Senior Network Security Design Engineer
Core4ce Careers
North Charleston, SC
The Senior Network Security Design Engineer serves as a technical subject matter expert responsible for designing, prototyping, documenting, and supporting enterprise network security…
Added 6 mins ago
AI Software Engineer, App Intelligence
Apple
Cupertino, CA
…resulting experience is good enough to trust. As a platform engineer, you will build the on-device software this depends on and make the resulting experiences reliable and fast. This is…
Added 6 mins ago
Mechatronics Engineer III
Allen Institute
Seattle, WA
Mechatronics Engineer III - Scientific Instrumentation & Process Engineering (SIPE)
Added 10 mins ago
TT
Senior Forward Deployed Engineer, Operate
True Talent Solutions Partners
New York, NY
- $170,000 to $205,000 USD / year
As a Senior Forward Deployed Engineer, you will lead technical delivery of AI agent deployments for healthcare and life sciences organizations. You will work directly with clients to build…
Added 12 mins ago
WB
Senior Software Engineer, AD/ADAS, Vehicle Platform
Woven by Toyota
Palo Alto, CA
- $140,000 to $230,000 USD / year
Added 13 mins ago
Senior Financial Planning & Analysis Manager
cyberu
United States
We're looking for a Senior Financial Planning & Analysis Manager
Added 14 mins ago
ZZ
Senior Software Development Engineer
ZINC Zillow, Inc.
Remote, United States
- $156,200 to $262,600 USD / year
The Senior Specialist, Software Engineering, Back End designs, builds, and operates scalable, reliable backend services and platforms that power Zillow’s consumer and partner experiences.…
Added 17 mins ago
FS
Senior Software Engineer- DevOps
Four Sea Group, Inc.
Aurora, CO
- $140,000 to $232,000 USD / year
Four Sea Group is seeking a Senior Software Engineer specializing in DevOps to define and advance secure, reliable, and repeatable software-delivery and infrastructure capabilities for…
Added 17 mins ago
FS
Senior Software Engineer – Mission Applications & Command and Control
Four Sea Group, Inc.
Aurora, CO
- $140,000 to $232,000 USD / year
Four Sea Group is seeking a Senior Software Engineer to design, develop, integrate, and advance full-stack software capabilities supporting complex command-and-control and mission systems.
Added 20 mins ago
L
Data Engineer Client Data Operations
Limetree
Remote, United States
- $115,000 to $145,000 USD / year
…execution of routine mail drops. You will work closely with our Senior Analytics Engineer, who depends on clean, consistent data for downstream analytics and reporting. This is an…
Added 29 mins ago
Senior Performance Analysis Engineer – Ground-based Missile Defense (MDA)
Mission Driven Research
Huntsville, AL
MDR is looking for an experienced, team-oriented Senior Performance Analysis Engineer to join our Huntsville, AL team. In this role you will support the analysis and assessment of…
Added 30 mins ago
Space Flight Dynamics Engineer
The Aerospace Corporation
El Segundo, CA · Chantilly, VA · Colorado Springs, CO · Albuquerque, NM
…simulation. We are looking for a Member of the Technical Staff/Senior Member of the Technical Staff to join the EOFDD team. The selected candidate will provide expertise for a wide range of…
Added 31 mins ago
Sr Leader, SDET
loanDepot
Irvine, CA · Plano, TX
…throughout the software development lifecycle. Serves as a senior technical advisor and recognized subject matter expert, driving adoption of modern quality engineering practices and…
Added 34 mins ago
Senior Silicon Photonics Automated Test Production Engineer
OpenLight Photonics
Santa Clara, CA
…experience in designing custom test hardware and programming in python. The position is onsite at our Santa Clara CA office and our San Jose CA…
Added 38 mins ago
Application Operations Analyst
VIATEQ Corporation
Washington, DC
…support activities. While escalating complex issues to the Senior Application Operations Engineer or Applications & Integration Lead as appropriate, this individual is expected to resolve…
Added 38 mins ago
Zscaler Integration Engineer - Mid
VIATEQ Corporation
Washington, DC
The ideal candidate is a technically proficient and security-focused cloud security engineer with solid, hands-on experience across key Zscaler platforms - including Zscaler Internet Access…
Added 38 mins ago
DevOps Cloud Engineer - EviCore - Remote
eviCore healthcare MSI, LLC
Remote, United States
As a Senior DevOps Engineer, you will interact with various functional groups such as Engineering, QA, Infrastructure, Security, Product Owners, Client Service, and Operations to ensure our…
Added 38 mins ago
IG
Hybrid (NYC) - Campus Network Engineer
Insight Global
San Jose, CA
- $150,000 to $160,000 USD / year
We are seeking an experienced Senior Campus Network Engineer with significant automation experience to design, deploy, migrate, and operate enterprise campus networks, with a strong focus…
Added 38 mins ago
IG
Sr. Network Engineer - WAN/MPLS - On site (Jersey City ,Palo Alto, Columbus, or Plano)
Insight Global
San Jose, CA
- $140,000 to $160,000 USD / year
We are seeking a highly experienced Senior Network Engineer with a strong background in Service Provider networking, WAN, MPLS, and IS-IS routing. The ideal candidate will have extensive…
Added 38 mins ago
IG
REMOTE- Sr Data Engineer/PM
Insight Global
Remote, United States
- $120,000 to $140,000 USD / year
Insight Global is seeking a Senior Data Engineer for a top healthcare client. This role is ideal for someone who combines strong hands-on data engineering experience with technical…
Added 40 mins ago
Senior IT Systems Engineer
10x Genomics
Pleasanton, CA
…innovation and productivity. We are looking for an exceptional Senior Systems Engineer with strong knowledge of Windows and virtualization platforms and extensive experience in building and…
Added 42 mins ago
S
Senior Data Scientist
SOSi
Huntsville, AL
- $83,164 to $172,725 USD / year
The ideal candidate will have a background in AI/ML model development and have experience in Python programming, handling SQL databases, and working in command line interfaces. You will…
Added 42 mins ago
Data Engineer
KBR Wyle Services, LLC
Beavercreek, OH
KBR is seeking a Senior Data Engineer (Lakehouse & Application Development) to design, develop, and optimize modern data platforms and web-based applications supporting Department of…
Added 44 mins ago
Senior Software Engineer
USA-Medtronic MiniMed, Inc 1017
Northridge, CA
We are seeking for a skilled and motivated Senior Software Embedded Engineer to join our dynamic team. The ideal candidate will have a strong background in embedded systems and software…
Added 47 mins ago
Senior Analytics Engineer
EnergyHub
Remote, United States
…transformation frameworks (e.g. dbt), and data science tools (e.g. Python/Jupyter, Julia,…

Addgene

Application Developer IV

Watertown, MA, United States

SALARY $130,000 - $145,000

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Job details

Pay: SALARY $130,000 - $145,000
Apply by: Feb 1, 2027
Posting language: English
Posted: Nov 3, 2026

Original job description

TITLE Application Developer IV (Full Time)

WHAT YOU WILL ACHIEVE

The Application Developer IV will join our Application Development team to architect and build features across our digital products and lead complex technical initiatives. You’ll help shape how we build and evolve the systems that power Addgene’s operations and researcher experience, so we can better serve researchers and partners with greater reliability, clarity, and speed as the organization grows.

POSITION REPORTS TO

Director of Application Development

APPROXIMATE START DATE

January 2027

SALARY $130,000 - $145,000

LOCATION & POLICY ON REMOTE WORK

This role is completely flexible to be fully remote, hybrid, or onsite, whatever works best for you! Our office is located at 490 Arsenal Way in Watertown, Massachusetts. Our beautiful office space has free, ample parking with a complimentary shuttle from Harvard Square and is on multiple bus lines, with the cost of public transit fully covered by Addgene (if onsite at least 2 days/week).

You must be a resident in one of the following states: MA, VT, ME, RI, NC, FL, CO and OH.

The Role:

The Application Development team builds and evolves the systems behind Addgene’s digital products as part of our digital product development practice, collaborating closely with Digital Technology teams and others across the organization, so researchers and partners can rely on the high-quality solutions we deliver. We are looking for a senior engineer who brings sound judgment to complex technical work, drives challenging initiatives forward while staying aligned with roadmap priorities, and helps the team adapt its engineering practices as AI and other emerging tools reshape how we build software.

To Succeed in this Role, You Will:

Lead complex digital product initiatives from discovery through delivery: turn prioritized opportunities into sound technical solutions, provide clear options and reliable cost estimates, and advance roadmap outcomes through to completion
Architect and deliver features across our digital products, balancing roadmap goals with long-term maintainability and paying down technical debt as you go
Build and evolve systems and workflows that maintain data integrity under concurrent use, retries, and partial failures
Help the team evaluate and adopt AI and other emerging tools in ways that improve how we build and deliver software
Improve our test strategy by choosing the right kinds of tests so the team can ship confidently and efficiently
Identify where performance improvements matter most and use data to guide and confirm the results
Help other engineers grow through mentoring, code review, and pair programming

Skills and Competencies Must-haves

6+ years building and maintaining production web applications, with deep Python and Django expertise
Working fluency across a modern web stack: JavaScript, HTML/CSS, SQL, REST APIs, Docker, Git and CI
A track record of shaping test strategy for a codebase, balancing speed, coverage and confidence, and helping others do the same
Experience designing systems that maintain data integrity under concurrent use, with sound transaction handling and idempotent operations
Strong instincts for clean, maintainable design, favoring explicit dependencies and well-scoped state over shortcuts
Deep experience with database performance in Django applications, from query design to indexing and profiling
Experience making significant data-model changes and migrations in a production application without disrupting users
Experience with search relevance, embeddings, or recommendation systems, including measuring whether results actually improved
Experience building, evaluating or integrating machine learning or LLM-based features, such as classification, ranking or features that work with scientific data

Behavioral competencies

Takes ownership of work from first idea through production, and follows through
Leads through influence: disagrees constructively, explains tradeoffs to non-engineers and builds shared approaches
Seeks and acts on feedback as readily as giving it
Mentors generously and shares knowledge openly
Thinks ahead about what could go wrong and raises risks early
Stays curious and keeps learning as tools and practices evolve

Nice to have (not required) None of these are necessary to apply. Each would be a welcome addition to the team.

Experience with AI-assisted development beyond individual use: helping define a team’s AI coding harness (shared agent instructions, guardrails, evaluations and automated quality checks) and keeping current as these tools and practices evolve
Designing versioned public APIs and role-based access control for external partners or customers
Advanced software design: layered or domain-driven design in Django, async or event-driven patterns, observability, and web security practices such as Content Security Policy
Third-party integrations and data pipelines, including responsible handling of personal data
Familiarity with molecular biology, lab workflows or research data

Addgene is an Equal Opportunity Employer and considers all applicants without regard to race, color, religion, national origin, sex, ancestry, sexual orientation, genetic information, gender identity, active military or veteran status, age, handicap, or any other characteristic protected by Federal, State or Local law.

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More about this job

Skills

Coaching
Python
Django
JavaScript
HTML
CSS
SQL
API development
Docker
Git
Machine learning
Statistical quality control
Molecular biology
Research

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