Jobs in the United States

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Added 1 min ago
Anticheat Engineer Intern - Summer 2027
Electronic Arts
Austin, TX
The Anticheat Engineer Intern will report to the Senior Anti-Cheat Engineer II.
Added 3 mins ago
Senior Principal Mechanical Engineer
INFOSYS NOVA HOLDINGS LLC
Boston, MA
Senior / Principal Mechanical Design Engineer – Medical Robotics
Added 4 mins ago
Sr Network Operations Engineer
loanDepot
Irvine, CA
…response, and crisis procedures; collaborates closely with Network Engineering and Cybersecurity teams on secure enterprise designs; and ensures duties are performed in accordance with…
Added 4 mins ago
Sr. Prototyping Engineer
Corning
Keller, TX
The Senior Prototyping Engineer designs, builds, and coordinates prototypes for optical connectivity solutions used in fiber optic networks. This hands-on role supports a low-volume,…
Added 4 mins ago
Prototyping Engineer
Corning
Keller, TX
The Prototyping Engineer helps design, build, and coordinate prototypes for optical connectivity solutions used in fiber optic networks. This is a hands-on role on a low-volume,…
Added 6 mins ago
U
Staff Software Engineer - Engineer
Uber
Sunnyvale, CA · San Francisco, CA · Seattle, WA
- $232,000 to $258,000 USD / year
Establish engineering excellence and operational best practices for mission-critical infrastructure, including architecture reviews, reliability standards, incident prevention, and…
Added 6 mins ago
U
Sr Software Engineer - Engineer
Uber
Sunnyvale, CA · San Francisco, CA · Seattle, WA
- $202,000 to $224,000 USD / year
Senior Software Engineer, Michelangelo Job Controller team
Added 7 mins ago
Production Engineer
Venturi
Huntsville, AL
Venturi, LLC A Chenega Company has an exciting career opportunity for a Production Engineer who will be directly responsible for verifying that critical missile flight hardware and…
Added 7 mins ago
AI Engineer
Coterie Applications Inc.
Remote, United States
The AI Engineer will work within the Research & Development (R&D) subdomain of Coterie’s broader Data and Analytics (DnA) department to build, test and deploy intelligent systems that…
Added 7 mins ago
C
Senior Site Reliability Engineer - (US-Remote)
ClickHouse
Remote, United States
- $150,000 to $230,000 USD / year
…secure services so we are building out our newly formed Site Reliability Engineering team. As one of the first joiners to our Reliability Engineering Team at ClickHouse, you will be…
Added 9 mins ago
Senior Software Engineer
Indeed
United States
As a Senior Software Engineer at Indeed, you will contribute to the development and evolution of software systems that support core products and services used by people around the world.…
Added 11 mins ago
Senior DevOps Engineer
Securiport
Reston, VA
We are seeking a skilled and motivated Senior (Sr.) DevOps Engineer to join our Release and DevOps team. In this role, you will work as a member in the development group, helping to…
Added 12 mins ago
Senior Software Engineer
TEXAS RESEARCH INSTITUTE AUSTIN INC
Austin, TX
TRI Austin is seeking an experienced Senior Software Engineer to lead the development, enhancement, and maintenance of software applications used to analyze, manage, and visualize complex…
Added 23 mins ago
Application Engineer C - Automotive Business Unit
NSK Americas
Ann Arbor, MI
…reconditioning, ball screw repair, product integration, application engineering, and predictive & preventative maintenance and reliability services. NSK's product and solutions are…
Added 23 mins ago
Senior Staff Software Engineer - Monetization
Nextdoor
Remote, United States
We are seeking a Senior Staff Software Engineer, reporting to the Head of Engineering, Monetization, to shape technical strategy and build scalable, reliable systems across Nextdoor’s…
Added 24 mins ago
Senior Developer Experience Engineer-AI Infrastructure Operations
Nscale
NY
This is a senior engineering role for someone who sets the bar for how Nscale ships software and then pulls the rest of the team up to it. You'll own the hardest problems on the delivery…
Added 24 mins ago
Senior Full-Stack Software Engineer (Arsenal 2)
Anduril Industries
Costa Mesa, CA
As a Senior Full Stack Software Engineer on the ArsenalOS Deployment initiative, you will build the full-stack web applications that power daily planning, work assignment, and team-lead…
Added 24 mins ago
Senior Backend Software Engineer (Arsenal2)
Anduril Industries
Costa Mesa, CA
As a Senior Full Stack Software Engineer on the Systems Integrity Group (SIG) initiative, you will build simulation and digital twin validation frameworks that verify manufactured output…
Added 25 mins ago
Senior Optical Transport Network Engineer
Comcast (CC) of Willow Grove
CO
The Senior Optical Transport Network Engineer is a critical technical leadership role responsible for the reliability, engineering, and evolution of Comcast's national optical backbone.…
Added 26 mins ago
K
Senior System Engineer
Kforce
Draper, UT
- $110,000 to $150,000 USD / year
Kforce has a client that is seeking a Senior Systems Engineer in Draper, UT.
Added 28 mins ago
Senior Data Integration Engineer
Everon
Irving, TX
The Senior Data Integration Engineer designs, develops, implements, and supports secure, scalable, and reliable integrations across enterprise applications, databases, cloud platforms,…
Added 30 mins ago
Senior CUAS Systems Engineer
Parsons Government Services Inc.
WV
Monitors system performance post-deployment and implements improvements to enhance reliability and effectiveness.
Added 31 mins ago
SA
Verification and Validation Engineer (R5430)
Shield AI
Dallas, TX
- $95,000 to $200,000 USD / year
As a Software Systems Test Engineer, you will be pivotal in driving integration testing of our full aircraft software system, integrating our autonomy, flight controller, and GCS software…
Added 34 mins ago
Launch and Test Engineer III (Avionics)
Blue Origin Personnel, LLC
Space Coast, FL
The Test and Launch Engineer will drive rate and reliability for New Glenn's critical launch infrastructure – including our massive Transporter/Erector, LC-36 Launch Pad, Mobile Test Stand,…
Added 34 mins ago
Senior Platform Engineer (Infrastructure)
Fleetio
Remote, United States
We're looking for a Senior Platform Engineer to join our growing Platform Engineering team. This role will have a direct impact on our customers, our engineers, and our business by ensuring…

Addgene

Application Developer IV

Watertown, MA, United States

SALARY $130,000 - $145,000

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Job details

Pay: SALARY $130,000 - $145,000
Apply by: Feb 1, 2027
Posting language: English
Posted: Nov 3, 2026

Original job description

TITLE Application Developer IV (Full Time)

WHAT YOU WILL ACHIEVE

The Application Developer IV will join our Application Development team to architect and build features across our digital products and lead complex technical initiatives. You’ll help shape how we build and evolve the systems that power Addgene’s operations and researcher experience, so we can better serve researchers and partners with greater reliability, clarity, and speed as the organization grows.

POSITION REPORTS TO

Director of Application Development

APPROXIMATE START DATE

January 2027

SALARY $130,000 - $145,000

LOCATION & POLICY ON REMOTE WORK

This role is completely flexible to be fully remote, hybrid, or onsite, whatever works best for you! Our office is located at 490 Arsenal Way in Watertown, Massachusetts. Our beautiful office space has free, ample parking with a complimentary shuttle from Harvard Square and is on multiple bus lines, with the cost of public transit fully covered by Addgene (if onsite at least 2 days/week).

You must be a resident in one of the following states: MA, VT, ME, RI, NC, FL, CO and OH.

The Role:

The Application Development team builds and evolves the systems behind Addgene’s digital products as part of our digital product development practice, collaborating closely with Digital Technology teams and others across the organization, so researchers and partners can rely on the high-quality solutions we deliver. We are looking for a senior engineer who brings sound judgment to complex technical work, drives challenging initiatives forward while staying aligned with roadmap priorities, and helps the team adapt its engineering practices as AI and other emerging tools reshape how we build software.

To Succeed in this Role, You Will:

Lead complex digital product initiatives from discovery through delivery: turn prioritized opportunities into sound technical solutions, provide clear options and reliable cost estimates, and advance roadmap outcomes through to completion
Architect and deliver features across our digital products, balancing roadmap goals with long-term maintainability and paying down technical debt as you go
Build and evolve systems and workflows that maintain data integrity under concurrent use, retries, and partial failures
Help the team evaluate and adopt AI and other emerging tools in ways that improve how we build and deliver software
Improve our test strategy by choosing the right kinds of tests so the team can ship confidently and efficiently
Identify where performance improvements matter most and use data to guide and confirm the results
Help other engineers grow through mentoring, code review, and pair programming

Skills and Competencies Must-haves

6+ years building and maintaining production web applications, with deep Python and Django expertise
Working fluency across a modern web stack: JavaScript, HTML/CSS, SQL, REST APIs, Docker, Git and CI
A track record of shaping test strategy for a codebase, balancing speed, coverage and confidence, and helping others do the same
Experience designing systems that maintain data integrity under concurrent use, with sound transaction handling and idempotent operations
Strong instincts for clean, maintainable design, favoring explicit dependencies and well-scoped state over shortcuts
Deep experience with database performance in Django applications, from query design to indexing and profiling
Experience making significant data-model changes and migrations in a production application without disrupting users
Experience with search relevance, embeddings, or recommendation systems, including measuring whether results actually improved
Experience building, evaluating or integrating machine learning or LLM-based features, such as classification, ranking or features that work with scientific data

Behavioral competencies

Takes ownership of work from first idea through production, and follows through
Leads through influence: disagrees constructively, explains tradeoffs to non-engineers and builds shared approaches
Seeks and acts on feedback as readily as giving it
Mentors generously and shares knowledge openly
Thinks ahead about what could go wrong and raises risks early
Stays curious and keeps learning as tools and practices evolve

Nice to have (not required) None of these are necessary to apply. Each would be a welcome addition to the team.

Experience with AI-assisted development beyond individual use: helping define a team’s AI coding harness (shared agent instructions, guardrails, evaluations and automated quality checks) and keeping current as these tools and practices evolve
Designing versioned public APIs and role-based access control for external partners or customers
Advanced software design: layered or domain-driven design in Django, async or event-driven patterns, observability, and web security practices such as Content Security Policy
Third-party integrations and data pipelines, including responsible handling of personal data
Familiarity with molecular biology, lab workflows or research data

Addgene is an Equal Opportunity Employer and considers all applicants without regard to race, color, religion, national origin, sex, ancestry, sexual orientation, genetic information, gender identity, active military or veteran status, age, handicap, or any other characteristic protected by Federal, State or Local law.

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More about this job

Skills

Coaching
Python
Django
JavaScript
HTML
CSS
SQL
API development
Docker
Git
Machine learning
Statistical quality control
Molecular biology
Research

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