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Added 3 days ago
Program Manager Intern
FlightSafety International Inc
Seattle, WA
This position is an individual contributor role reporting to Senior Director, Cloud and Production Engineering.
Added 5 days ago
Director, Sales Engineering
Mattermost
United States
We are seeking an experienced Director of Sales Engineering to build and lead Mattermost's global technical presales team.
Added 5 days ago
Application Developer IV
Addgene
Watertown, MA
…the high-quality solutions we deliver. We are looking for a senior engineer who brings sound judgment to complex technical work, drives challenging initiatives forward while staying aligned…
Added 6 days ago
Senior Manager, AI & Platform Engineering
Procore Technologies, Inc.
TX
We are looking for a Senior Manager, AI & Platform Engineering to lead a newly formed team that runs Procore’s enterprise AI platform (Claude) and the business/IT side of the Atlassian…
Added 6 days ago
Senior Program Manager
General Dynamics Mission Systems, Inc
Taunton, MA
General Dynamics Mission Systems has an immediate opening for a Senior Program Manager within the Network and Communication Systems Line of Business. The position provides an opportunity to…
Added 9 days ago
Sr. systems analyst- Microsoft365
Starbucks Coffee Company
Nashville
As a Senior Systems Analyst – Collaboration Engineering, you will help shape and support the technology platforms that enable partners around the world to communicate, collaborate, share…
Added 9 days ago
Sr. systems analyst- Microsoft365
Starbucks Coffee Company
TN
As a Senior Systems Analyst – Collaboration Engineering, you will help shape and support the technology platforms that enable partners around the world to communicate, collaborate, share…
Added 9 days ago
Lead IT Program Manager
Klaviyo
Boston, MA
…help the teams around you do the same. You will report to the Director of Corporate IT and partner with IT leaders, Engineering, Security, Finance, Procurement, Legal, and Business…
Added 9 days ago
H
Senior Infrastructure Engineer
Harvey
Remote, United States
- $153,000 to $210,500 USD / year
…layer that every Harvey IT system runs on. You'll join Core IT Engineering & Infrastructure, reporting to Dave Vega, Director of IT, alongside engineers who own identity and collaboration,…
Added 9 days ago
H
Senior Infrastructure Engineer
Harvey
New York, NY
- $153,000 to $210,500 USD / year
…layer that every Harvey IT system runs on. You'll join Core IT Engineering & Infrastructure, reporting to Dave Vega, Director of IT, alongside engineers who own identity and collaboration,…
Added 11 days ago
Director, PE/VC & Portfolio Partnerships
ZoomInfo Technologies LLC
Waltham, MA
…Success, and Solutions Partners, coordinating with Product and Engineering to meet quality, security, and timeline…
Added 11 days ago
Manager, Device Mgmt Team, Amazon Devices Beta Team
Amazon.com Services LLC
Seattle, WA
…will also maintain a regular weekly reporting cadence with senior leadership. The ability to synthesize complex operational data into clear, actionable narratives for director-level and…
Added 13 days ago
Senior Program Coordinator
Saronic Technologies
Austin, TX
The Senior Program Coordinator partners cross-functionally with engineering, test, and mission stakeholders to remove blockers and align on priorities, owns the systems used to track…
Added 13 days ago
Senior Director of Product Marketing
Pindrop
Remote, United States
As the Senior Director of Product Marketing at Pindrop, you’ll play a crucial role at the nexus of Marketing, Product, and Sales, with substantial impact potential on company growth. Your…
Added 14 days ago
Executive Assistant (Hybrid, Sunnyvale)
CrowdStrike, Inc.
Sunnyvale, CA
2+ years of executive administrative support experience, including direct support of senior leaders (Director level or above), or equivalent combination of experience and education
Added 16 days ago
Sr, IT Administrator
Zones LLC.
Washington DC, DC
…management and resource allocation processes. In partnership with Director of IT and looking across all of the programs and projects the team is working on, you will implement and develop…
Added 17 days ago
S
VP of Engineering
SiteZeus
Remote, United States
- $150,000 to $170,000 USD / year
SiteZeus is hiring a hands-on VP of Engineering to lead a small, senior product engineering team operating AI-first. You will own the technical direction, production quality, and delivery…
Added 17 days ago
Senior IT Engineer (Onsite)
Axon
Boston, MA
As Axon’s Senior Client Engineer, you will be a senior technical owner within the IT Engineering team, helping shape how Axon’s global workforce connects, collaborates, and stays secure.…
Added 18 days ago
IG
Director of Architecture
Insight Global
Chicago, IL
- $190,000 USD / year
Partner with Engineering and architecture leaders on promotion gates, source control, environments, CI/CD, security, and production standards
Added 18 days ago
Director of Product Management, Platform
Cobalt
Remote, United States
Cobalt is hiring a Director of Product Management, Platform to own the product surfaces that connect our pentesting offerings to the rest of a customer's security and business stack, and…
Added 22 days ago
UO
Director of Operations - Office of Academic Services - Penn Engineering
University of Pennsylvania
Philadelphia, PA
- $67,155 to $105,000 USD / year
Reporting to the Managing Director of the Office of Academic Services (OAS), the Director of Operations is an agile, outcomes-driven leader who advances operational excellence and service…
Added 23 days ago
Mobile Product Owner
MTech Systems (MTech Systems USA)
United States
…development tooling, training, and ongoing opportunities to work with senior and executive…
Added 23 days ago
Financial Product Owner
MTech Systems (MTech Systems USA)
Atlanta, GA
…development tooling, training, and ongoing opportunities to work with senior and executive…
Added 23 days ago
H
Privacy Associate
Handshake
San Francisco, CA
- $120,000 to $149,000 USD / year
…spot-check fulfillment across connected systems, and partner with Engineering to resolve integration issues. Track request volume and turnaround times, and recommend workflow…
Added 23 days ago
Director, Data Architecture (AI Enablement)
Digital Asset
NY
AI is only as useful as the data it can safely reach. As Director, Data Architecture, you will own how our organization’s information, from databases to data warehouses to email, documents,…

Addgene

Application Developer IV

Watertown, MA, United States

SALARY $130,000 - $145,000

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Job details

Pay: SALARY $130,000 - $145,000
Apply by: Feb 1, 2027
Posting language: English
Posted: Nov 3, 2026

Original job description

TITLE Application Developer IV (Full Time)

WHAT YOU WILL ACHIEVE

The Application Developer IV will join our Application Development team to architect and build features across our digital products and lead complex technical initiatives. You’ll help shape how we build and evolve the systems that power Addgene’s operations and researcher experience, so we can better serve researchers and partners with greater reliability, clarity, and speed as the organization grows.

POSITION REPORTS TO

Director of Application Development

APPROXIMATE START DATE

January 2027

SALARY $130,000 - $145,000

LOCATION & POLICY ON REMOTE WORK

This role is completely flexible to be fully remote, hybrid, or onsite, whatever works best for you! Our office is located at 490 Arsenal Way in Watertown, Massachusetts. Our beautiful office space has free, ample parking with a complimentary shuttle from Harvard Square and is on multiple bus lines, with the cost of public transit fully covered by Addgene (if onsite at least 2 days/week).

You must be a resident in one of the following states: MA, VT, ME, RI, NC, FL, CO and OH.

The Role:

The Application Development team builds and evolves the systems behind Addgene’s digital products as part of our digital product development practice, collaborating closely with Digital Technology teams and others across the organization, so researchers and partners can rely on the high-quality solutions we deliver. We are looking for a senior engineer who brings sound judgment to complex technical work, drives challenging initiatives forward while staying aligned with roadmap priorities, and helps the team adapt its engineering practices as AI and other emerging tools reshape how we build software.

To Succeed in this Role, You Will:

Lead complex digital product initiatives from discovery through delivery: turn prioritized opportunities into sound technical solutions, provide clear options and reliable cost estimates, and advance roadmap outcomes through to completion
Architect and deliver features across our digital products, balancing roadmap goals with long-term maintainability and paying down technical debt as you go
Build and evolve systems and workflows that maintain data integrity under concurrent use, retries, and partial failures
Help the team evaluate and adopt AI and other emerging tools in ways that improve how we build and deliver software
Improve our test strategy by choosing the right kinds of tests so the team can ship confidently and efficiently
Identify where performance improvements matter most and use data to guide and confirm the results
Help other engineers grow through mentoring, code review, and pair programming

Skills and Competencies Must-haves

6+ years building and maintaining production web applications, with deep Python and Django expertise
Working fluency across a modern web stack: JavaScript, HTML/CSS, SQL, REST APIs, Docker, Git and CI
A track record of shaping test strategy for a codebase, balancing speed, coverage and confidence, and helping others do the same
Experience designing systems that maintain data integrity under concurrent use, with sound transaction handling and idempotent operations
Strong instincts for clean, maintainable design, favoring explicit dependencies and well-scoped state over shortcuts
Deep experience with database performance in Django applications, from query design to indexing and profiling
Experience making significant data-model changes and migrations in a production application without disrupting users
Experience with search relevance, embeddings, or recommendation systems, including measuring whether results actually improved
Experience building, evaluating or integrating machine learning or LLM-based features, such as classification, ranking or features that work with scientific data

Behavioral competencies

Takes ownership of work from first idea through production, and follows through
Leads through influence: disagrees constructively, explains tradeoffs to non-engineers and builds shared approaches
Seeks and acts on feedback as readily as giving it
Mentors generously and shares knowledge openly
Thinks ahead about what could go wrong and raises risks early
Stays curious and keeps learning as tools and practices evolve

Nice to have (not required) None of these are necessary to apply. Each would be a welcome addition to the team.

Experience with AI-assisted development beyond individual use: helping define a team’s AI coding harness (shared agent instructions, guardrails, evaluations and automated quality checks) and keeping current as these tools and practices evolve
Designing versioned public APIs and role-based access control for external partners or customers
Advanced software design: layered or domain-driven design in Django, async or event-driven patterns, observability, and web security practices such as Content Security Policy
Third-party integrations and data pipelines, including responsible handling of personal data
Familiarity with molecular biology, lab workflows or research data

Benefits

At Addgene, we prioritize our people with comprehensive benefits that support both your work and your life.

Full-time employees receive:

85% company-paid healthcare premiums including dental (extending to family members) and 100% company-paid deductible
Life, short-term, and long-term disability insurance.
22 days of PTO on top of our winter break closure and company holidays. ‌
A 403(b) retirement account with a company match of 4% after 2 years of employment. ‌
Annual professional development stipend
Annual lifestyle stipend
Fully paid public transportation for employees who work onsite two or more days per week. ‌
Onsite yoga classes & wellness reimbursement
Pet insurance
Annual financial planning one-on-ones with our plan administrator

The Organization:

Addgene is a self-sustaining, thriving nonprofit life sciences organization that was created to help scientists share DNA-based research materials called plasmids. ‌Our mission is to accelerate research and discovery by improving access to useful research materials and information. Since its founding in 2004, Addgene has grown and innovated in order to assist the research community by archiving and sharing useful DNA-based research materials and resources (such as plasmids, viral vectors, and recombinant antibodies) with scientists worldwide. Broad availability of these unique research materials promotes the advancement of research in a wide variety of disciplines, including cancer, heart disease, and neurodegenerative disorders, while increasing collaboration within the scientific community. Learn more here from our President & Executive Director, Chonnettia Jones. As an “Addgenie”, you will work collaboratively with internal teams to help fulfill this mission, including scientists, software engineers and product developers, renowned customer support teams, scientific content creators, and business development professionals. Together, Addgenies create and enhance opportunities for research scientists to deposit their materials, find products and resources easily on our website, and receive unparalleled support in receiving and expertly using our materials in their own experiments.

Life at Addgene:

Addgene is a truly special place to work. We are an inclusive and welcoming group of people from many backgrounds and perspectives. We believe in flexibility, actively supporting professional development, and a strong work/life balance. We value sharing ideas and feedback with each other and the company. Learn more about Life at Addgene We also believe it's important to make time for fun, whether it's eating popcorn at our outdoor movie night, sharing cute photos in the #pets slack channel, or sampling whiskeys at our end-of-year social hour. Bookworms, athletes, gamers, parents, foodies, and more will find community in Addgene's events and Slack channels. Addgene works with a global scientific community in which the best innovations and discoveries come from diverse teams. We continuously learn from and celebrate a variety of perspectives and seek to create an environment where all people can feel heard, respected, and valued.

Addgene is an Equal Opportunity Employer and considers all applicants without regard to race, color, religion, national origin, sex, ancestry, sexual orientation, genetic information, gender identity, active military or veteran status, age, handicap, or any other characteristic protected by Federal, State or Local law.

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Skills

Coaching
Python
Django
JavaScript
HTML
CSS
SQL
API development
Docker
Git
Machine learning
Statistical quality control
Molecular biology
Research

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