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Senior Cybersecurity Engineer
Tau Six
Chantilly, VA
…cybersecurity and system integration services to the US National Security market. We have an immediate need for a Senior Cybersecurity Engineer and are seeking a motivated self-starter with…
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Welding Engineer
National Steel and Shipbuilding Com
San Diego, CA
…qualifications. Reports to the position of the Senior Welding Engineer/Chief Weld Engineer/Manager of Welding Engineer. Maintains internal contact with fitting and Welding Production areas…
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Tech Accelerate Graduate Program - Software Engineer I
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Alpharetta, GA
…gain hands-on experience in big data, AI, SRE, DevOps, and security engineering. They also get exposure to leading cloud platforms like AWS and Azure. The program focuses on technical and…
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Technical Director
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- $170,000 to $195,000 USD / year
…engineering-internal development and bench testing, borrowing technicians when engineer-directed test or assembly work needs them. You'll bring manufacturability and documentation quality…
Added 2 mins ago
Senior Network Engineer (Active Secret Clearance)
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Co-lead secure network integration design with the security engineer and review high- and low-level designs, network diagrams, and future-state documentation.
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Software Engineer Data & AI II (Full Time) – United States
Cisco
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…AI, helping to transform how Cisco and its customers harness information and intelligent…
Added 3 mins ago
Material Flow Engineer
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The Material Flow Engineer supports Yamaha Motor Manufacturing Corporation of America’s Material Flow Engineering activities by developing and improving material flow processes that support…
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Where required by state law and/or city ordinance; this employer will provide the Social Security Administration (SSA) and, if necessary, the Department of Homeland Security (DHS), with…
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Staff Electrical Engineer, Avionics
Longshot
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- $180,000 to $220,000 USD / year
We are hiring a Staff Electrical Engineer to own electrical engineering across Longshot, including the people, the architecture and the hardware. You will provide leadership and expertise…
Added 6 mins ago
Senior Quality Engineer
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Must be a U.S. citizen. Applicants selected may be subject to a government security investigation and must meet eligibility requirements for access to classified information.
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Presales Solution Engineer - Edge AI
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The Presales Solution Engineer – Edge AI is a customer-facing technical leader responsible for transforming Edge AI opportunities into measurable business outcomes and production-ready…
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Senior Developer Experience Engineer-AI Infrastructure Operations
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Software Engineer, Imaging
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Senior Full-Stack Software Engineer (Arsenal 2)
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MDP operates at the boundary between Operational Technology and Information Technology. Our systems live where factory-floor machines, edge compute, and OT networks meet enterprise…
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Senior Backend Software Engineer (Arsenal2)
Anduril Industries
Costa Mesa, CA
MDP operates at the boundary between Operational Technology and Information Technology. Our systems live where factory-floor machines, edge compute, and OT networks meet enterprise…
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Software Engineer III - Data Engineering | Infrastructure Platforms
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As a Software Engineer III at JPMorganChase within the Infrastructure Platforms organization, you serve as a seasoned member of an agile team to design and deliver trusted market-leading…
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Regulatory Engineer
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…reliable communication for satellite missions such as national security, global connectivity, and disaster response. With a vertically integrated approach, Northwood designs, builds, and…
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Incoming Quality Technician
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Software Development Engineer, AI Foundations, Amazon Leo Networking Software
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Senior System Engineer
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- $110,000 to $150,000 USD / year
Senior System Engineer will take ownership of complex escalations involving servers, storage, networking, security, and operational platforms
Added 8 mins ago
Senior Security Engineer - Cloud
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Muon seeks a Senior Security Engineer to join our Security Engineering & IT team. The ideal candidate brings deep expertise across multiple security domains and can operate independently to…
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Senior Security Engineer - Aerospace
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Engineer I - ST - Partner Technology (Nashville, TN - U.S.)
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Now Brewing: engineer I - Front-end iOS! #tobeapartner
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Endpoint Support Specialist
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The Endpoint Support Specialist is responsible for the deployment, configuration, maintenance, security, and support of Matheny's end-user technology environment. This includes desktop and…
Added 9 mins ago
Domain Consultant - Public Sector
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Addgene

Application Developer IV

Watertown, MA, United States

SALARY $130,000 - $145,000

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Job details

Pay: SALARY $130,000 - $145,000
Apply by: Feb 1, 2027
Posting language: English
Posted: Nov 3, 2026

Original job description

TITLE Application Developer IV (Full Time)

WHAT YOU WILL ACHIEVE

The Application Developer IV will join our Application Development team to architect and build features across our digital products and lead complex technical initiatives. You’ll help shape how we build and evolve the systems that power Addgene’s operations and researcher experience, so we can better serve researchers and partners with greater reliability, clarity, and speed as the organization grows.

POSITION REPORTS TO

Director of Application Development

APPROXIMATE START DATE

January 2027

SALARY $130,000 - $145,000

LOCATION & POLICY ON REMOTE WORK

This role is completely flexible to be fully remote, hybrid, or onsite, whatever works best for you! Our office is located at 490 Arsenal Way in Watertown, Massachusetts. Our beautiful office space has free, ample parking with a complimentary shuttle from Harvard Square and is on multiple bus lines, with the cost of public transit fully covered by Addgene (if onsite at least 2 days/week).

You must be a resident in one of the following states: MA, VT, ME, RI, NC, FL, CO and OH.

The Role:

The Application Development team builds and evolves the systems behind Addgene’s digital products as part of our digital product development practice, collaborating closely with Digital Technology teams and others across the organization, so researchers and partners can rely on the high-quality solutions we deliver. We are looking for a senior engineer who brings sound judgment to complex technical work, drives challenging initiatives forward while staying aligned with roadmap priorities, and helps the team adapt its engineering practices as AI and other emerging tools reshape how we build software.

To Succeed in this Role, You Will:

Lead complex digital product initiatives from discovery through delivery: turn prioritized opportunities into sound technical solutions, provide clear options and reliable cost estimates, and advance roadmap outcomes through to completion
Architect and deliver features across our digital products, balancing roadmap goals with long-term maintainability and paying down technical debt as you go
Build and evolve systems and workflows that maintain data integrity under concurrent use, retries, and partial failures
Help the team evaluate and adopt AI and other emerging tools in ways that improve how we build and deliver software
Improve our test strategy by choosing the right kinds of tests so the team can ship confidently and efficiently
Identify where performance improvements matter most and use data to guide and confirm the results
Help other engineers grow through mentoring, code review, and pair programming

Skills and Competencies Must-haves

6+ years building and maintaining production web applications, with deep Python and Django expertise
Working fluency across a modern web stack: JavaScript, HTML/CSS, SQL, REST APIs, Docker, Git and CI
A track record of shaping test strategy for a codebase, balancing speed, coverage and confidence, and helping others do the same
Experience designing systems that maintain data integrity under concurrent use, with sound transaction handling and idempotent operations
Strong instincts for clean, maintainable design, favoring explicit dependencies and well-scoped state over shortcuts
Deep experience with database performance in Django applications, from query design to indexing and profiling
Experience making significant data-model changes and migrations in a production application without disrupting users
Experience with search relevance, embeddings, or recommendation systems, including measuring whether results actually improved
Experience building, evaluating or integrating machine learning or LLM-based features, such as classification, ranking or features that work with scientific data

Behavioral competencies

Takes ownership of work from first idea through production, and follows through
Leads through influence: disagrees constructively, explains tradeoffs to non-engineers and builds shared approaches
Seeks and acts on feedback as readily as giving it
Mentors generously and shares knowledge openly
Thinks ahead about what could go wrong and raises risks early
Stays curious and keeps learning as tools and practices evolve

Nice to have (not required) None of these are necessary to apply. Each would be a welcome addition to the team.

Experience with AI-assisted development beyond individual use: helping define a team’s AI coding harness (shared agent instructions, guardrails, evaluations and automated quality checks) and keeping current as these tools and practices evolve
Designing versioned public APIs and role-based access control for external partners or customers
Advanced software design: layered or domain-driven design in Django, async or event-driven patterns, observability, and web security practices such as Content Security Policy
Third-party integrations and data pipelines, including responsible handling of personal data
Familiarity with molecular biology, lab workflows or research data

Addgene is an Equal Opportunity Employer and considers all applicants without regard to race, color, religion, national origin, sex, ancestry, sexual orientation, genetic information, gender identity, active military or veteran status, age, handicap, or any other characteristic protected by Federal, State or Local law.

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Skills

Coaching
Python
Django
JavaScript
HTML
CSS
SQL
API development
Docker
Git
Machine learning
Statistical quality control
Molecular biology
Research

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