Jobs in the United States

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Added 8 days ago
Cloud Engineer III
Quality Technology Services, LLC
Remote, United States
The Cloud Engineer III is primarily responsible for deploying, configuring, troubleshooting and ensuring continuity of client and QTS physical and virtualized infrastructures and operating…
Added 8 days ago
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Research Engineer
Twenty
Arlington, VA · New York, NY
- $159,000 to $195,000 USD / year
Twenty builds offensive cyber capabilities for the United States and its allies. We’re looking for a Research Engineer who can turn an understanding of adversary behavior into software that…
Added 8 days ago
Senior Software Engineer, GRC
Reddit
Remote, United States
We’re hiring a GRC Software Engineer to build and improve the software that supports Reddit’s security and compliance programs. You will design, implement, and maintain integrations, data…
Added 8 days ago
System Test Engineer
Picarro, Inc
Santa Clara, CA
Picarro is looking for a System Test Engineer to join our Operations/Manufacturing team. You will work directly with sophisticated analytical instruments, performing system-level testing,…
Added 8 days ago
Research Engineer (Electrical / Power Systems) - 1704
SARA Inc
Colorado Springs, CO
…an exceptionally talented, motivated, and creative Research Engineer to support rapid Research & Development (R&D) and power systems research activities out of our facility in Colorado…
Added 8 days ago
Software Engineer II
Samsara
Remote, United States
As a Software Engineer II on this team, you will own customer-facing features from concept to production. You will partner closely with Product, Design, and ML and AI engineering teams to…
Added 8 days ago
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Solution Engineer
Snowflake
Remote, United States
- $138,000 to $181,125 USD / year
Location Requirement: Must be based in Delaware, Maryland, or Virignia
Added 8 days ago
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Account Engineer
Snowflake
Menlo Park, CA · Seattle, WA
- $138,000 to $181,125 USD / year
Location: Candidate must be based in or near Seattle or Menlo Park
Added 8 days ago
Software Validation Engineer, Energy Storage
Redwood Materials
San Francisco, CA
As a Software Validation Engineer, you are the bridge between pure code and physical reality. You will design and maintain the environments - Software-in-the-Loop (SIL) and…
Added 8 days ago
Test Engineer
Itron, Inc.
United States of America, SC
As a Test Engineer, you will contribute to the testing and validation of electricity meter firmware and supporting software. You will participate in the automated firmware testing…
Added 8 days ago
Senior Vice President, Full-Stack Engineer
BNY
Jersey City, NJ · NY
Job holder must have understanding interdependencies and business impact of future IT plans · Job holder must have broad experience with multi-platform development tools and toolkits.
Added 8 days ago
Hypersonic Aerothermal Test Engineer/Analyst
Torch Technologies, Inc.
United States
Torch Technologies is seeking a highly skilled Hypersonic Aerothermal Test Engineer/Analyst to support the 715th Test Squadron (715 TS) at the Arnold Engineering Development Complex (AEDC).…
Added 8 days ago
Mid Level SDET
Vidoori
Hyattsville
Must have experience with Playwright & TypeScript
Added 8 days ago
Sr. Mechanical Engineer
Intuitive
Sunnyvale, CA
Intuitive Surgical designs and manufactures highly complex mechanical systems for use in surgery. The Mechanical Engineer is responsible for developing, documenting, and validating new…
Added 8 days ago
Senior Test Engineer
Ralliant
Hollister, CA
Pacific Scientific EMC is expanding and seeking a highly skilled Senior Test Engineer to lead the development, automation, and qualification of electrical test systems that support new and…
Added 8 days ago
Manufacturing Systems Engineer – Digital Manufacturing
Fairbanks Morse Defense
Walpole, MA
The Digital Manufacturing Engineer – Manufacturing Data & Automation is responsible for developing and implementing data-driven manufacturing solutions across machining, assembly,…
Added 8 days ago
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Integration Engineer
TEKsystems
Palm Beach, FL
- $125,000 to $135,000 USD / year
Added 8 days ago
Software Engineer - Humanoid Sim-to-Real
Apptronik
Austin, TX
…help solve the ever-increasing labor shortage. As a Controls Engineer – Actuation on our Controls team, you will own both the actuator models that make our simulation trustworthy and the…
Added 8 days ago
QA Engineer 3
Onto Innovation Inc-US
United States
We are looking for an experienced and skilled Software Test Engineer to join our team. The ideal candidate will have a strong background in verification and validation of complex software.…
Added 8 days ago
Senior Software Engineer
Onto Innovation Inc-US
United States
We are looking for a Software Engineer to join the Inspection Business Unit (IBU). Whether you are launching your career or bringing years of deep expertise, what matters most is how you…
Added 8 days ago
Software Engineer II - Electronic Trading Services - Automation Software Engineer
JPMC page
Jersey City, NJ
Design, develop, test, deploy, maintain, and improve applications supporting network automation initiatives.
Added 8 days ago
Analyst Lead, Cybersecurity
FIS INC (OLD FIS)
United States
As a Senior Application Security Engineer in FIS Cybersecurity, you will be a hands-on technical leader in our Attack Surface Management function.
Added 8 days ago
Developer Experience (DevEx) Engineer
Williams International
Pontiac, MI
Must have strong analytical and critical thinking skills
Added 8 days ago
Application Developer IV
Addgene
Watertown, MA
Fully paid public transportation for employees who work onsite two or more days per week
Added 8 days ago
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Senior SIEM Data Engineer
SSB&T
Quincy, MA
- $120,000 to $202,500 USD / year
We are looking for a Senior SIEM Data Engineer reporting directly to the Cyber Data Engineering Manager. You will lead the design, onboarding, transformation, validation, and operational…

Addgene

Application Developer IV

Watertown, MA, United States

SALARY $130,000 - $145,000

Apply on addgene.applytojob.com About employer

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Job details

Pay: SALARY $130,000 - $145,000
Apply by: Feb 1, 2027
Posting language: English
Posted: Nov 3, 2026

Original job description

TITLE Application Developer IV (Full Time)

WHAT YOU WILL ACHIEVE

The Application Developer IV will join our Application Development team to architect and build features across our digital products and lead complex technical initiatives. You’ll help shape how we build and evolve the systems that power Addgene’s operations and researcher experience, so we can better serve researchers and partners with greater reliability, clarity, and speed as the organization grows.

POSITION REPORTS TO

Director of Application Development

APPROXIMATE START DATE

January 2027

SALARY $130,000 - $145,000

LOCATION & POLICY ON REMOTE WORK

This role is completely flexible to be fully remote, hybrid, or onsite, whatever works best for you! Our office is located at 490 Arsenal Way in Watertown, Massachusetts. Our beautiful office space has free, ample parking with a complimentary shuttle from Harvard Square and is on multiple bus lines, with the cost of public transit fully covered by Addgene (if onsite at least 2 days/week).

You must be a resident in one of the following states: MA, VT, ME, RI, NC, FL, CO and OH.

The Role:

The Application Development team builds and evolves the systems behind Addgene’s digital products as part of our digital product development practice, collaborating closely with Digital Technology teams and others across the organization, so researchers and partners can rely on the high-quality solutions we deliver. We are looking for a senior engineer who brings sound judgment to complex technical work, drives challenging initiatives forward while staying aligned with roadmap priorities, and helps the team adapt its engineering practices as AI and other emerging tools reshape how we build software.

To Succeed in this Role, You Will:

Lead complex digital product initiatives from discovery through delivery: turn prioritized opportunities into sound technical solutions, provide clear options and reliable cost estimates, and advance roadmap outcomes through to completion
Architect and deliver features across our digital products, balancing roadmap goals with long-term maintainability and paying down technical debt as you go
Build and evolve systems and workflows that maintain data integrity under concurrent use, retries, and partial failures
Help the team evaluate and adopt AI and other emerging tools in ways that improve how we build and deliver software
Improve our test strategy by choosing the right kinds of tests so the team can ship confidently and efficiently
Identify where performance improvements matter most and use data to guide and confirm the results
Help other engineers grow through mentoring, code review, and pair programming

Skills and Competencies Must-haves

6+ years building and maintaining production web applications, with deep Python and Django expertise
Working fluency across a modern web stack: JavaScript, HTML/CSS, SQL, REST APIs, Docker, Git and CI
A track record of shaping test strategy for a codebase, balancing speed, coverage and confidence, and helping others do the same
Experience designing systems that maintain data integrity under concurrent use, with sound transaction handling and idempotent operations
Strong instincts for clean, maintainable design, favoring explicit dependencies and well-scoped state over shortcuts
Deep experience with database performance in Django applications, from query design to indexing and profiling
Experience making significant data-model changes and migrations in a production application without disrupting users
Experience with search relevance, embeddings, or recommendation systems, including measuring whether results actually improved
Experience building, evaluating or integrating machine learning or LLM-based features, such as classification, ranking or features that work with scientific data

Behavioral competencies

Takes ownership of work from first idea through production, and follows through
Leads through influence: disagrees constructively, explains tradeoffs to non-engineers and builds shared approaches
Seeks and acts on feedback as readily as giving it
Mentors generously and shares knowledge openly
Thinks ahead about what could go wrong and raises risks early
Stays curious and keeps learning as tools and practices evolve

Nice to have (not required) None of these are necessary to apply. Each would be a welcome addition to the team.

Experience with AI-assisted development beyond individual use: helping define a team’s AI coding harness (shared agent instructions, guardrails, evaluations and automated quality checks) and keeping current as these tools and practices evolve
Designing versioned public APIs and role-based access control for external partners or customers
Advanced software design: layered or domain-driven design in Django, async or event-driven patterns, observability, and web security practices such as Content Security Policy
Third-party integrations and data pipelines, including responsible handling of personal data
Familiarity with molecular biology, lab workflows or research data

Benefits

At Addgene, we prioritize our people with comprehensive benefits that support both your work and your life.

Full-time employees receive:

85% company-paid healthcare premiums including dental (extending to family members) and 100% company-paid deductible
Life, short-term, and long-term disability insurance.
22 days of PTO on top of our winter break closure and company holidays. ‌
A 403(b) retirement account with a company match of 4% after 2 years of employment. ‌
Annual professional development stipend
Annual lifestyle stipend
Fully paid public transportation for employees who work onsite two or more days per week. ‌
Onsite yoga classes & wellness reimbursement
Pet insurance
Annual financial planning one-on-ones with our plan administrator

The Organization:

Addgene is a self-sustaining, thriving nonprofit life sciences organization that was created to help scientists share DNA-based research materials called plasmids. ‌Our mission is to accelerate research and discovery by improving access to useful research materials and information. Since its founding in 2004, Addgene has grown and innovated in order to assist the research community by archiving and sharing useful DNA-based research materials and resources (such as plasmids, viral vectors, and recombinant antibodies) with scientists worldwide. Broad availability of these unique research materials promotes the advancement of research in a wide variety of disciplines, including cancer, heart disease, and neurodegenerative disorders, while increasing collaboration within the scientific community. Learn more here from our President & Executive Director, Chonnettia Jones. As an “Addgenie”, you will work collaboratively with internal teams to help fulfill this mission, including scientists, software engineers and product developers, renowned customer support teams, scientific content creators, and business development professionals. Together, Addgenies create and enhance opportunities for research scientists to deposit their materials, find products and resources easily on our website, and receive unparalleled support in receiving and expertly using our materials in their own experiments.

Life at Addgene:

Addgene is a truly special place to work. We are an inclusive and welcoming group of people from many backgrounds and perspectives. We believe in flexibility, actively supporting professional development, and a strong work/life balance. We value sharing ideas and feedback with each other and the company. Learn more about Life at Addgene We also believe it's important to make time for fun, whether it's eating popcorn at our outdoor movie night, sharing cute photos in the #pets slack channel, or sampling whiskeys at our end-of-year social hour. Bookworms, athletes, gamers, parents, foodies, and more will find community in Addgene's events and Slack channels. Addgene works with a global scientific community in which the best innovations and discoveries come from diverse teams. We continuously learn from and celebrate a variety of perspectives and seek to create an environment where all people can feel heard, respected, and valued.

Addgene is an Equal Opportunity Employer and considers all applicants without regard to race, color, religion, national origin, sex, ancestry, sexual orientation, genetic information, gender identity, active military or veteran status, age, handicap, or any other characteristic protected by Federal, State or Local law.

Apply on addgene.applytojob.com About employer

More about this job

Skills

Coaching
Python
Django
JavaScript
HTML
CSS
SQL
API development
Docker
Git
Machine learning
Statistical quality control
Molecular biology
Research

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Work requirements

Jobs in the United States